Okami Amaterasu, the origin of all that is good and mother to all of Nippon, finds herself sent on a strange mission in another world. With only a few friends at her side, she embarks for this new land, only to find it devoid of all human life. Worse still, there is a usurper of her title of sun goddess.

What's a goddess to do, when your noy companions are a hyperactive puppy and a phoenix with anger issues?

Meanwhile, the moon and sun in Equestria are going out of control, raising and lowering when they feel like it. And a powerful magical presence has been detected- a goddess. Celestia and Luna are worried, and call in the Elements.