Having gotten a few months of rest and relaxation since her last adventure, Lyra is asked to head out to the mysterious Island of Echidna to speak with its ruler, Princess Medusa. Joining her is lovable former treasure hunter Ditzy Doo, geologist Maud Pie, and the mysterious Ocean Wave, a thouroughly unique mare.

However, disaster strikes Lyra again, and she and her three companions end up on the shores of Lemuria, a land once home to the first true empire in the world.

And the inhabitants are still there