Fortuna has returned, Pharynx is healing from his injuries, their family is whole once more, everything is as it should be. Until Fortuna brings up the fact that Thorax and Pharynx's father Metamorphosis, a changeling they'd assumed long gone, was in fact still alive. In Equestria. And she wants to bring him home too. Naturally, both brothers have some feelings about this, good and bad. As tensions rise between the family, Fortuna sets them on one last journey: to return their father, her cousin, to the Hive. But she will find not everyone is quite so eager to quicken this reunion, nor will they all be joyful upon the inevitable meeting.

After all, how else can they be expected to react when faced with the presence of a father they never knew?